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    If you do, you will have an email fprtuna and password for your account. A Philosophy of Nature. Start reading El mito de la diosa Fortuna on your Kindle in under a minute. Furthermore, we analyze the chronology and dynamics of the GABI, the evolution of some South American mammalian groups through time, and the Quaternary mammalian extinctions.

    The geology and paleogeography of Central America and northern South America have been intensively described e. After different stages, Pangea split into the supercontinents of Gondwana and Laurasia Rapela and Pankhurst ; Iturralde-Vinent Rifting processes since Jurassic times provoked the separation of Western and Eastern Gondwana. The first evidence of separation of Africa and North America is the magnetic anomaly of Ma earliest Middle Jurassic; Benedetto The Author s A.

    Cione et al. According to the fossil record, a biotic interchange of land vertebrates e. In the Panamanian Isthmus region, Late Cretaceous to Early Middle Miocene beds of the San Blas Complex and the Clarita, Darien, and Porcona formations were deposited in deep depths in an open ocean, low energy, essentially non-siliciclastic sedimentary environment distant from South America.

    However, the oldest land bridge between West Indies and the main land dates to the latest Eocene earliest Oligocene according to the Gaarlandia hypothesis and bridged the Greater Antilles with South America through the Aves Ridge Jestrow et al. After this, and until the latest Cenozoic, South America was practically an island continent. The gray arrow indicates complete interchange of Atlantic and Pacific waters.

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    This is a postcollisional phase as the Panama microplate and the area become extensively emerged. Strongly modified from Coates et al. Land exposed by lower than today s positions of the Pleistocene sea level during maximum glacial advance is indicated.

    Black areas are elevations above m a. Modified from Woodburne 2. The opening of the Caribbean was associated with large displacement faults such as the Volcanic Trans-Mexican Alignment.

    Along the Pacific Ocean, the subduction of the Farallon plate generated a volcanic arc that was the origin of the Greater Antilles Benedetto Neritic depths were predominant throughout the Darien 5 2. Besides, extensive emergence and rapid uplift in the Central American Isthmus in the latest Miocene is suggested by the absence of Pliocene deposits from either the Darien or the Panama Canal Basin and of sediments younger than 5 Ma in the Atrato Basin of Colombia Coates et al.

    Central American archipelago putatively served as a sweepstakes route during the Late Miocene when there was a limited biotic interchange among North and South America Brown and Lamolino Complete docking and widespread uplift of the Central American Isthmus including Panama is reflected by a widespread unconformity at about Ma.

    This could have fostered the entry of the Procyonidae to South America Woodburne Shallow marine conditions adjacent to emergent lands between the latest Miocene and the Early Pliocene from Costa Rica to Colombia and in Colombia and in Costa Rica in the Pliocene are indicated by marine beds.

    Subsequent to this, the region was further strongly uplifted, including the final emergence of the Panamanian district Woodburne Fig.

    Woodburne has suggested that land mammals began to extend their ranges across Central America soon after the initial tectonic closure from about 12 Ma. The first sloths arrived in North America at about 9 Ma, that is only a minimum age regarding their actual crossing of the isthmian region, and whether they were present in Central America, prior to their occurrence in continental North America, remains to be determined. In fact, the degree to which Central America acted as a holding pen for any trans-isthmian dispersal is still an open question Woodburne The fossil evidence indicates that the main interchange began at about 2.

    Iturralde-Vinent suggests that the isthmus suffered interruptions shown by interchange record known between the Pacific and Caribbean seas e. The interruption became definitive at about 0.

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    Since then, the Caribbean geography is similar to the present one and new marine endemic species emerged. However, the Central American region increased its area during the sea lowstands during glaciations Fig. More recently, a new model of formation of the Isthmus of Panama proposes that its establishment is much older 15 Ma; Coates and Stallard The closure at 15 Ma was suggested because by this time the alignment of the blocks leaves no space for trans-isthmian marine passages.

    Besides, Prothero et al. This connection would permit the first interchange see below.

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    Finally, Prothero et al. However, considerable debate is currently in progress in the geological community see also Leigh et al. Supporting the first scenario, molecular data from cross-isthmian divergence times of marine organisms reported on 34 trans-isthmian sister clades of fish, mollusks, and crustaceans suggest the absence of a marine dispersal barrier between the Atlantic and Pacific until the Late Neogene Baker et al. Later, Charles R. Darwin called them the Pampean formation Tonni The Spanish use of the Pampean Formation outlasted and gave rise to others such as Pampiano Formation or Pampean sediments, the latter commonly used in geological and stratigraphic literature since the end of the nineteenth century Fidalgo et al.

    Auguste Bravard was the first researcher to give a brief mineralogical description of these deposits which he called indistinctly Pampa Formation, Pampa silt, or Pampa terrain see Tonni He recognized their mainly eolian origin, in contrast to what was supported by d Orbigny and Darwin see Tonni and Pasquali ; Tonni et al.

    This classification consisted of fourteen geological horizons or stages including, among others, the Araucana, Pampeana, Tehuelche, Querandina, and Ariana formations Doering The stratigraphic classification of Doering was the base for that one proposed by Florentino Ameghino. Thus, Ameghino divided these sediments into terreno pampeano inferior, terreno pampeano superior, and terreno pampeano lacustre. In his publication on the stratigraphy and paleontology of the Atlantic Bonaerian coast Fig.

    The name ensenadan was created by Ameghino on the basis of the observations made in the town of Ensenada, Buenos Aires Province, during the construction of the harbor of La Plata see Soibelzon et al. When Ameghino consolidated his stratigraphic framework, in the last decade of the nineteenth and early twentieth centuries, he used the terms formation and stage horizon or subformation which currently define units of the lithostratigraphic and chronostratigraphic classification.

    However, at the time of 8 16 2 Continental Relationships, Chronostratigraphy, Climates Ameghino, the meaning of these terms was different. A formation was a unit defined from the fossil content, where the lithology was not the essential element or the only one in the definition. The stage or subformation or geological horizon was merely a subdivision of the formation and did not differ largely from the current biostratigraphic units see Cione and Tonni ; Tonni In the mid-twentieth century, and based on previous observations, the Italian naturalist, Gioacchino Frenguelli , continued with the stratigraphic framework of Ameghino, but simplified apparently with another connotation, adjusted to the new stratigraphic classifications established since the s see Tonni Frenguelli s stratigraphic framework is crystallized in his contribution of ; he recognized Series and a set of stages subordinate to them, i.

    However, Frenguelli , never explained that his stratigraphic framework responded to formal categories.